Scientific publications

European Redlist of Freshwater Fishes

Freshwater Fishes of North Africa and the Middle East Freyhof, J. & Brooks, E. 2011. European Red List of Freshwater Fishes. Luxembourg: Publications Office of the European Union, 61 pp.

Handbook of European Freshwater Fishes

Handbook of European Freshwater FishesKottelat, M. & Freyhof, J. 2007. Handbook of European Freshwater Fishes. Kottelat, Cornol and Freyhof, Berlin, xiv + 646 pp.

Freshwater Fishes of North Africa and the Middle East

Freshwater Fishes of North Africa and the Middle East Freyhof, J., et al. Freshwater fishes of North Africa and the Middle East. (In prep)

Scientific publications

(Selected, since 2000); (Impact factor)



Denys, G. P. J., Persat, H., Dettai, A., Geiger, M. F., Freyhof, J., Fesquet, J. & Keith, P. (2018) Genetic and morphological discrimination of three species of ninespined stickleback Pungitius spp. (Teleostei, Gasterosteidae) in France with the revalidation of Pungitius vulgaris (Mauduyt, 1848). Journal of Zoological Systematics and Evolutionary Research, 56, 77–101. (2.444)


Freyhof, J., Özulug, M., Kaya, C. Baycelebi, C., & Turan, D. 2018. Redescription of Alburnus kotschyi Steindachner, 1863, with comments on Alburnus sellal adanensis Battalgazi, 1944 (Teleostei: Leuciscidae). Zootaxa, 4382 (3), 573-582. (0.972)


Friedrich, T., Wiesner, C., Zangl, L., Daill, D., Freyhof, J., & Korblmüller, S. 2018. Romanogobio skywalkeri, a new gudgeon (Teleostei: Gobionidae) from the upper Mur River, Austria. Zootaxa, 4403 (2), 336-350. (0.972)

  Pichler, A. Ahnelt, H., Kirchner, S., Sattmann, H., Haring, E., Handschuh, S., Freyhof, J., Victor, R. & Kruckenhauser, L. 2018. The morphological diversity of Garra barreimiae (Teleostei: Cyprinidae). Environmental Biology of Fishes, (1.255)

Freyhof, J., Kaya, C., Bayçelebi, E., Geiger, M. & Turan, D. 2018. Generic assignment of Leuciscus kurui Bogutskaya from the upper Tigris drainage, and a replacement name for Alburnus kurui Mangit & Yerli (Teleostei: Leuciscidae). Zootaxa, 4410 (1): 113–135. (0.972)


  Carrizo, S., Lengyel, S., Kapusi, F., Szabolcs, M., Kasperidus, H., Scholz, M., Markovic, D., Freyhof, J., Cid, N., Cardoso, A.-C., & Darwall, W. 2017. Critical catchments for freshwater biodiversity conservation in Europe: identification, prioritisation and gap-analysis. Journal of Applied Ecology, 54, 4, 1209-1218. (5.196)

Freyhof, J. & Abdullah, Y.S. 2017. Two new species of Oxynoemacheilus from the Tigris drainage in Iraqi Kurdistan (Teleostei: Nemacheilidae). Zootaxa, 4238, 73-87. (0.972)

  Freyhof, J. & Özulug, M. 2017. Oxynoemacheilus hazarensis, a new species from Lake Hazar in Turkey, with remarks on O. euphraticus (Teleostei: Nemacheilidae). Zootaxa, 4247, 378-390. (0.982)
  Behrmann-Godel, J., Nolte, A.W., Kreiselmaier, J., Berka, R. &Freyhof, J. 2017. The first European cave fish. Current Biology 27, R257-R258. (8.851)
  Freyhof, J. Kaya, C. & Turan, D. 2017. Oxynoemacheilus kentritensis, a new species from the upper Tigris drainage in Turkey with remarks on O. frenatus (Teleostei: Nemacheilidae). Zootaxa, 4258, 551-560. (0.972)
  Freyhof, J. & Geiger, M. 2017. Oxynoemacheilus zarzianus, a new loach from the Lesser Zab River drainage in Iraqi Kurdistan (Teleostei: Nemacheilidae). Zootaxa, 4273, 258-270. (0.972)
  Tutman, P., Freyhof, J. Dulčić, J. Glamuzina, B. & Geiger, M. 2017. Lampetra soljani, a new brook lamprey from the southern Adriatic Sea basin (Petromyzontiformes: Petromyzontidae). Zootaxa, 4273, 531-548. (0.972)
  Brahimi, A. Freyhof, J., Henrard, A. & Libois, R. 2017. Luciobarbus chelifensis and L. mascarensis, two new species from Algeria (Teleostei: Cyprinidae). Zootaxa, 4277, 32-50. (0.972)
  Sommerwerk, N., Wolter, C., Freyhof, J. & Tockner, K. 2017. Components and drivers of change in European freshwater fish faunas. Journal of Biogeography (4.248)
  Freyhof, J., Özulug, M. & Sac, G. 2017. Neotype designation of Aphanius iconii, first reviser action to stabilise the usage of A. fontinalis and A. meridionalis and comments on the family group names of fishes placed in Cyprinodontidae (Teleostei: Cyprinodontiformes). Zootaxa, 4294, 5, 573-585. (0.972)
  Buj, I., Marcic, Z., Caleta, M., Sanda, R., Geiger, M. F., Freyhof, J., Machordom, A., Vukic, J. 2017. Ancient connections among the European rivers and watersheds revealed from the evolutionary history of the genus Telestes (Actinopterygii; Cypriniformes). PLoS One, 12, 12, e0187366 (3.54)
  Mohammadian-Kalat, T., Esmaeili, H. R., Aliabadia, M. & Freyhof, J. 2017. Re-description of Alburnus doriae, with comments on the taxonomic status of A. amirkabiri. A. mossulensis, A. sellal and Petroleuciscus esfahani (Teleostei: Cyprinidae). Zootaxa. 4323, 4, 487-502. (0.972)
  Freyhof, J., Weissenbacher, A. & Geiger, M. 2017. Aphanius kruppi, a new killifish from Oman with comments on the A. dispar species group (Cyprinodontiformes: Aphaniidae). Zootaxa, 4338, 3, 557-573. (0.972)
  Carrizo, S. F., Jähnig, S., C., Bremerich; V., Freyhof, J., Harrison, I., He, F., Langhans, S. D., Tockner, K., Zarfl, C. & Darwall, W. 2017. Freshwater megafauna: flagships for freshwater biodiversity under threat. BioScience, 67, 10, 919-927. (5.378)


Freyhof, J., M. F. Geiger, K. Golzarianpour& Patimar, R. 2016. Sasanidus, a new generic name for Noemacheilus kermanshahensis Banarescu & Nalbant, with discussion of Ilamnemacheilus and Schistura (Teleostei; Nemacheilidae). Zootaxa, 4107: 65-80 (0.994)

Mousavi-Sabet, H., Ganjbakhsh, B., Geiger, M. F. & Freyhof, J. 2016. Redescription of Gobio nigrescens from the Hari River drainage (Teleostei: Cyprinidae). Zootaxa, 4114:71-80. (0.994)


Zareian, H., Esmaeili, H. R. & Freyhof, J. 2016. Capoeta anamisensis, a new species from the Minab and Hasan Langhi River drainages in Iran (Teleostei: Cyprinidae). Zootaxa, 4083 (1), 126-142. (0.994)


Hashemzadeh Segherloo, I., Ghaedrahmati, N. & Freyhof, J. 2016. Eidinemacheilus, a new generic name for Noemacheilus smithi Greenwood (Teleostei; Nemacheilidae). Zootaxa, 4147 (4): 466–476. (0.994)


Hashemzadeh Segherloo, I., Abdoli, A., Eagderi, S., Esmaeili, H. R., Sayyadzadeh, G., Bernatchez, L., Hallerman, E., Geiger, M. F., Özulug, M., Laroche, J. & Freyhof, J. 2016. Dressing down: convergent reduction of the mental disc in Garra (Teleostei: Cyprinidae) in the Middle East. Hydrobiologia, DOI 10.1007/s10750-016-2902-8 (2.051)

  Lyon, R. G., Geiger, M. F. & Freyhof, J. 2016. Garra sindhi, a new species from the Jebel Samhan Nature Reserve in Oman (Teleostei: Cyprinidae). Zootaxa, 4154 (1): 079–088. (0.994)

Turak, E., Harrison, I., Dudgeon, D., Abell, R., Bush, A., Darwall, D., Finlayson, C. M., Ferrier, S., Freyhof, J., Hermoso V., Juffe-Bignoli, D., Linke, S., Nel, J., Patricio, H. C., Pittock, J., Raghavan, R., Revenga, C., Simaika, J. P. & DeWever, A. 2016. Essential Biodiversity Variables for measuring change in global freshwater biodiversity. Biological Conservation, (3.985)

  Khaefi, R., Esmaeili, H. R., Sayyadzadeh, G., Geiger, M. F. & Freyhof, J. 2016. Squalius namak, a new chub from Lake Namak basin in Iran (Teleostei: Cyprinidae). Zootaxa 4169(1):145. (0,994)

Freyhof, J. 2016. Oxynoemacheilus karunensis, a new species from the Persian Gulf basin Teleostei: Nemacheilidae). Zootaxa, 4175(1): 94-100. (0.994)


Freyhof, J. 2016. Redescription of Garra elegans (Günther, 1868), a poorly known species from the Tigris River drainage (Teleostei: Cyprinidae). Zootaxa, 4173(5): 496-500. (0.994)


Freyhof, J., Abdullah, Y.S., Ibrahim, H., Ararat, K. & Geiger, M. 2016. Eidinemacheilus proudlovei, a new subterranean loach from Iraqi Kurdistan (Teleostei; Nemacheilidae). Zootaxa, 4173(3): 225-236. (0.994)


Turak, E., Dudgeon, D., Harrison, I. J., Freyhof, J. De Wever, A., Revenga, C., Garcia-Moreno, J., Abell, R., Culp, J. M., Lento, J., Mora, B. Hilarides, L. & Flink, S. (2016) Observations of Inland Water Biodiversity: Progress, Needs and Priorities. Pp. 165-186. In: Walters, M. & Scholes, R.J. (Eds). 2016. The GEO Handbook on Biodiversity Observation Networks. Springer, 326 pp.




Mousavi-Sabet, H., S. Vatandoust,H. R. Esmaeili, M. F. Geiger & J. Freyhof. 2015. Cobitis avicennae, a new species of spined loach from the Tigris River drainge (Teleostei: Cobitidae). Zootaxa, 3914 (5): 558–568. (1.648)


Piria, M., R. Rajeev & J. Freyhof. 2015. Editorial 2015: Efforts towards better quality in the Croatian Journal of Fisheries (Ribarstvo). Croatian Journal of Fisheries, 73: 1-5.


Yang, L., T. Sado, M. V. Hirt, E. Pasco-Viel, M. Arunachalam, J. Li, X. Wang, J. Freyhof, K. Saitoh, A. M. Simons, M. Miya, S. He & R. L Mayden. 2015. Phylogeny and polyploidy: Resolving the classification of cyprinine fishes (Teleostei: Cypriniformes). Molecular Phylogenetics and Evolution, 85: 97-116. (4.02)


Mousavi-Sabet, H., G. Sayyadzadeh, H. R. Esmaeili, S. Eagderi, R. Patimar & J. Freyhof. 2015. Paracobitis hircanica, a new crested loach from the southern Caspian Sea basin (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 24: 339-346. (1.01)


Dümpelmann, C, & J. Freyhof. 2015. First record of the fathead minnow Pimephales promelas Rafinesque, 1820 in Germany. Lauterbornia, 79: 173-180.


Freyhof J., G. Sayyadzadeh, H. R. Esmaeili & M. Geiger, 2015. Review of the genus Paraschistura from Iran with description of six new species (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 26:1-48. (1.01)


Harrison, I., García, N., Freyhof, J., Neubert, E., Amr, Z., Van Damme, D., & Cumberlidge, N. 2015. Chapter 1. Background. pp. 1-6. In: García, N., Harrison, I., Cox, N. and Tognelli, M.F. (compilers). (2015). The Status and Distribution of Freshwater Biodiversity in the Arabian Peninsula. Gland, Switzerland, Cambridge, UK and Arlington, USA: IUCN.

  Freyhof, J., Hamidan, N. A., Feulner, G. R. & Harrison, I. 2015. Chapter 3. The Status and Distribution of Freshwater Fishes of the Arabian Peninsula pp. 16-29. In: García, N., Harrison, I., Cox, N. and Tognelli, M.F. (compilers). (2015). The Status and Distribution of Freshwater Biodiversity in the Arabian Peninsula. Gland, Switzerland, Cambridge, UK and Arlington, USA: IUCN.

Kissling, D., A. Hardisty, E. Alonso Garcia, M. Santamaria, F. De Leo, G. Pesole, J. Freyhof, D. Manset, S. Wissel, J. Konijn & W. Los. 2015. Towards global interoperability for supporting biodiversity research on essential biodiversity variables (EBVs). Biodiversity, DOI: 10.1080/14888386.2015.1068709


Sayyadzadeh, G., · H. R. Esmaeili &· J. Freyhof . 2015.  Garra mondica, a new species from the Mond River drainage with remarks on the genus Garra from the Persian Gulf basin in Iran (Teleostei: Cyprinidae). Zootaxa, 4048:75-89. DOI:10.11646/zootaxa.4048.1.4. (0.91)


 Behrens-Chapuis,· S., F. Herder, · H. R. Esmaeili,· J. Freyhof, · N. A. Hamidan,· M. Özuluğ,· R. Šanda· & M. F. Geiger. 2015. Adding nuclear rhodopsin data where mitochondrial COI indicates discrepancies – can this marker help to explain conflicts in cyprinids? DNA Barcodes, 3: 187-199.


Darwall, W. & J. Freyhof. 2015. Lost fishes, who is counting? The extend of threat to freshwater fish biodiversity. Pp 1-35. In: Closs G. P., M. Krkosek & J. D. Olden (eds.) Conservation Of Freshwater Fishes.Cambridge University Press.



Geiger, M., F. Herder, M. Monaghan, V. Almada, R. Barbieri, M. Bariche, P. Berrebi, J. Bohlen, M. Casal-Lopez, G. Delmastro, G. Denys, A. Dettai, I. Doadrio, E. Kalogianni, H. Kärst, M. Kottelat, M. Kovacic, M. Laporte, M. Lorenzoni, Z. Marcic, M. Özulug, A. Perdices, S. Perea, H. Persat, S. Porcellotti, C. Puzzi, J. Robalo, R. Sanda, M. Schneider, V. Slechtova, M. Stumboudi, S. Walter, & J. Freyhof. 2014. Spatial heterogeneity in the Mediterranean Biodiversity Hotspot affects barcoding accuracy of its freshwater fishes. Molecular Ecology Resources, doi: 10.1111/1755-0998.12257 (7.432)


Esmaeili, H. R., Sayyadzadeh, G., Özuluğ, M., Geiger, M. & Freyhof, J. 2014. Three new species of Turcinoemacheilus from Iran and Turkey (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 24: 257-273. (1.52)


Freyhof, J., H. Kärst & M. Geiger. 2014. Valencia robertae, a new killifish from lower Pinios River, Greece (Cyprinodontiformes: Valenciidae). Ichthyological Exploration of Freshwaters, 24: 289-298. (1.52)


Markovic, D., S. Carrizo, J. Freyhof, N. Cid, L. Szabolcs, M. Scholz, H. Kasperdius & W. Darwall. 2014. Europe’s freshwater biodiversity under climate change: distribution shifts and conservation needs. Diversity and Distributions, DOI: 10.1111/ddi.12232. (6.122)


Marčić, Z., D. Mrdak, D. Milošević, P. Simonović, M. Piria, B. Kalamujić, T. Tomljanović, M. Povž, A. Adrović, S. Šljuka, M. Mrakovčić, A. Duplić, B. Hrašovec, M. Ivanc, D. Zabric, S. Weiss & J. Freyhof. 2014. Halting the loss of biodiversity – the Huchen in the Danube. Croatian Journal of Fisheries, 72: 138-139


Freyhof; J. & Özuluğ, M. 2014. Acanthobrama thisbeae, a new species of bream from southern Anatolia, Turkey (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 25: 1-10. (1.52)


Freyhof, J., H. R. Esmaeili, G. Sayyadzadeh & M. Geiger. Review of the crested loaches of the genus Paracobitis from Iran and Iraq with the description of four new species (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 25: 11-38. (2.275)


Hamidan, N. A., M.. F. Geiger & J. Freyhof. 2014. Garra jordanica, a new species from the Dead Sea basin with remarks on the relationship of G. ghorensis, G. tibanica and G. rufa (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 25: 223-236. (1.52)


Esmaeili, H. R., H. Mousavi-Sabet, G. Sayyadzadeh, S. Vatandoust & J. Freyhof. 2014. Paracobitis atrakensis, a new species of crested loach from northeastern Iran (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 25: 237-242. (1.52


Darwall, W., S. Carrizo, C. Numa, V. Barrios, J. Freyhof & K. Smith. 2014. Freshwater Key Biodiversity Areas in the Mediterranean Basin Hotspot. Infoming species conservation and development planning in freshwater ecosystems. IUCN, Cambridge, UK and Malaga, Spain.


Freyhof, J. F. G. Ekmekçi, A. Ali, N. R. Khamees, M. Özuluğ, N. Hamidan, F. Küçük & K.. G. Smith. 2014. Chapter 3. Freshwater fishes, 19-42. In: Smith, K. G., V. Barrios, W. R. T. Darwall & C. Numa  ](Editors). 2014. The status and distribution of freshwater biodiversity in the Eastern Mediterranean. Cambridge, UK, Malaga, Spain and Gland, Switzerland: IUCN. xiv+132pp.