Scientific publications

European Redlist of Freshwater Fishes

Freshwater Fishes of North Africa and the Middle East Freyhof, J. & Brooks, E. 2011. European Red List of Freshwater Fishes. Luxembourg: Publications Office of the European Union, 61 pp.

Handbook of European Freshwater Fishes

Handbook of European Freshwater FishesKottelat, M. & Freyhof, J. 2007. Handbook of European Freshwater Fishes. Kottelat, Cornol and Freyhof, Berlin, xiv + 646 pp.

Freshwater Fishes of North Africa and the Middle East

Freshwater Fishes of North Africa and the Middle East Freyhof, J., et al. Freshwater fishes of North Africa and the Middle East. (In prep)

Scientific publications

(Selected, since 2000); (Impact factor)


  Symonová, R., Majtánová, Z., Sember A., Staaks, G. B. O., Bohlen, J., Freyhof, J., Rábová, M. & Ráb, P. 2013. Genome differentiation in a species pair of coregonine fishes: an extremely rapid speciation driven by stress-activated retrotransposons mediating extensive ribosomal DNA multiplications. BMC Evolutionary Biology, 13:42. DOI: 10.1186/1471-2148-13-42. (3.521)
  Pereira, H. M., Ferrier, S., Walters, M., Geller, G., Jongman R. H. G., Scholes, R. J., Bruford, M., Brummitt, N., Butchart, S. H. M., Cardoso, A., Coops, N. C., Dulloo, E., Faith, D. P., Freyhof, J., Gregory, R. D., Heip, C., Höft, R., Hurtt, G., Jetz, W., Karp, D., McGeoch, M. A., Obura, D., Onoda, Y., Pettorelli, N., Reyers, B., Sayre, R., Scharlemann, J. P.W., Stuart, S. N., Turak, E., Walpole, M. & Wegmann, M. 2013. Essential biodiversity variables for global earth observation. Science, 339: 277-278. (31.201)

 Esmaeili, H. R., Sayyadzadeh, G., Ghasemian, S., Mirghiyasi, S., Zamaneian, N. R. & Freyhof, J. Length-weight relationships of Turcinoemacheilus kosswigi Bănărescu & Nalbant, 1964 (Teleostei:Nemacheilidae) in southwestern Iran and its relation to habitat structure. Journal of Applied Ichthyology, 29: 290-291. (0.945)


Filipe, A. F., Markovic, D., Pletterbauer, F., Tisseuil, C., De Wever, A., Schmutz, S., Bonada, N. & Freyhof J. 2013. Forecasting fish distribution along stream networks: brown trout (Salmo trutta) in Europe. Diversity and Distributions, (DOI: 10.11, 1–13. doi:10.1111/ddi.12086) (4.830)


Esmaeili, H. R., Malekzehi, H. Pazira, A. & Freyhof, J. 2013. First record of the Kalabans, Bangana dero (Hamilton, 1822) from Iran (Teleostei: Cyprinidae). Zoology of the Middle East, 59: 89-91. (0.49)


Golzarianpour, K., Abdoli, A., Patimar, P. & Freyhof, J. 2013. Turcinoemacheilus hafezi, a new loach from the Zagroz Mountains, Iran (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 24: 41-48. (2.20)


Elp, M., Özuluğ, M., Şen, S. & Freyhof, J. 2013. Validation of Alburnus timarensis from the Lake Van basin, eastern Anatolia (Teleostei: Cyprinidae). Zoology in the Middle East, 59: 235-244. (0.49)


  Markovic, D., Freyhof, J., Wolter, C. 2012. Where are all the fish: Potential of biogeographical maps to project current and future distribution patterns of freshwater species. PLoS ONE 7(7): e40530. doi:10.1371/journal.pone.0040530 (4.411)
  Levin, B. A., Freyhof, J., Lajbner, Z., Perea, S., Abdoli, A., Gaffaroglu, M., Özulug, M., Roubenyan, H. R. & Salnikov V. B. 2012. Phylogenetic relationships of the algae scraping cyprinid genus Capoeta (Teleostei: Cyprinidae). Molecular Phylogenetics and Evolution, 62: 542–549. (3.889)
   Yang, L., Arunachalam, M., Sado, T., Levin, B. A., Golubtsov, A. S., Freyhof, J., Friel, J. P., Chen, W.-J., Hirt, M. V., Manickam, R., Agnew, M. K., Simons, A. M., Saitoh, K., Miya, M., Mayden, R. L., & S. He, 2012. Molecular phylogeny of the cyprinid tribe Labeonini (Teleostei: Cypriniformes). Molecular Phylogenetics and Evolution, (3.609)



  Mehner, T., Freyhof, J. & Reichard, M. 2011. Summary and Perspective on Evolutionary Ecology of Fishes. Evolutionary Ecology, 25: 547–556. (2.398)
  Freyhof; J. 2011. Diversity and distribution of freshwater gobies from the Mediterranean, the Black and Caspian Seas, pp. 279-288. In: Patzner R. A., Van Tassell, J. L. Kovačić M. & Kapoor, B. G. (Eds). The Biology of Gobies. Science Publishers, Enfield, New Hampshire, 652 pp.
  Esmaeili, H. R., Gholamifard, A. & Freyhof J. 2011. Ichthyofauna of Zarivar Lake (Iran) with the first records of Hemiculter leucisculus and Alburnus hohenackeri in the Tigris drainage. Electronic Journal of Ichthyology, 7: 1-6.
  Özuluğ, M. & Freyhof; J. 2011. Review of the genus Squalius in Western and Central Anatolia, with the description of four new species (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 22: 107-148. (0.828)
  Harrison, I. J., Chakrabarty, P., Freyhof, J. & J. F. Craig 2011. Correct nomenclature and recommendations for preserving and cataloguing voucher material and genetic sequences. Journal of Fish Biology, 78: 1283-1290. (1.330)
  Golzarianpour, K., Abdoli, A. & Freyhof, J. 2011. Oxynoemacheilus kiabii, a new loach from Karkheh drainage, Iran (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 22: 201-208. (0.828)
  Freyhof, J., Erk'akan, F., Özeren, C. & Perdices, A. J. 2011. Remarks on the western Palearctic loach genus Oxynoemacheilus (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 22: 301-312. (0.828)


  Hoffmann, M., …., Freyhof, J., …., & Stuart, S. N. 2010. The impact of conservation on the status of the world’s vertebrates. Science, 330, 10: 1503-1509. (29.747)
  Mehner, T., Pohlmann, K., Elkin, C., Monaghan, M., Nitz, B. & Freyhof, J. 2010. Genetic population structure of sympatric and allopatric populations of Baltic ciscoes (Coregonus albula complex, Teleostei, Coregonidae). BMC Evolutionary Biology, 10: 85. (4.294)
  Perea, S., Bohme, M., Zupancic, P., Freyhof, J., Sanda, R., Özuluğ, M., Abdoli, A., & Doadrio, I. 2010. Phylogenetic relationships and biogeographical patterns in Circum-Mediterranean Subfamily Leuciscinae (Teleostei, Cyprinidae) inferred from both mitochondrial and nuclear data. BMC Evolutionary Biology, 10: 265. (4.294)
  Tang, K. L., Agnew, M. K., Chen, W.-J, Hirt, M. V., Sado, T., Schneider, L. M., Freyhof, J., Sulaiman, Z., Swartz, E., Vidthayanon, C., Saitoh, M. M. K., Simons, A. M., Wood, R. M. & Mayden, R. L. 2010. Systematics of the subfamily Danioninae (Teleostei: Cypriniformes: Cyprinidae). Molecular Phylogenetics and Evolution, 57: 189-214. (3.556)
  Symonová, R., Pelikánová, Š., Staaks, G. B.O., Freyhof, J. & Ráb, P. 2010. Cytogenetic study of Baltic (Coregonus albula) and Stechlin (C. fontanae) ciscoes species pair from Lake Stechlin, Germany: another test of Qumsiyeh’s hypothesis. Pp. 753-754. In: Słota, E. (Ed). 19th International Colloquium on animal cytogenetics and gene mapping. Chromosome Research, 18: 711–755. (3.230)
  Lewin, W.-C., Freyhof, J., Huckstorf, V., Mehner, T. & Wolter, C. 2010. When no catches matter: coping with zeros in environmental assessments. Ecological Indicators, 10: 572-583. (3.102)
  Mehner, T., Busch, S., Helland, I.P., Emmrich, M. & Freyhof, J. 2010. Temperature-related nocturnal vertical segregation of coexisting coregonids. Ecology of Freshwater Fish, 19: 408-419. (1.512)
  Pereira, A. M., Robalo, J. I., Freyhof, J., Maia, C., Fonseca, J. P., Valente, A. & Almada, V. C. 2010. Phylogeographical analysis reveals multiple conservation units in brook lampreys of Portuguese streams. Journal of Fish Biology, 77: 361–371. (1.226)
  Freyhof, J. & Özuluğ, M. 2010. Pseudophoxinus hittitorum, a new species of spring minnow from Central Anatolia (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 21: 239-245. (0.940)


  Mehner, T., Pohlmann, K., Elkin, C., Monaghan, M. & Freyhof, J. 2009. Genetic mixing from enhancement stocking in commercially exploited vendace populations. Journal of Applied Ecology, 46: 1340-1349. (4.220)
  Almada, V. C., Robalo, J. I., Levy, A., Freyhof, J., Bernardi, G., & Doadrio, I. 2009. Phylogenetic analysis of Peri-Mediterranean blennies of the genus Salaria: molecular insight on the colonization of freshwaters. Molecular Phylogenetics and Evolution, 52: 424-431. (3.994)
  Larmuseau, M. H. D., Freyhof, J., Volckaert, F. A. M. & Van Houdt, J. K. J. 2009. Matrilinear phylogeography and demographical patterns of the freshwater fish Rutilus rutilus: implications for taxonomy and conservation. Journal of Fish Biology, 75: 332-253. (1.404)
  Helland, I. P., Vollestad, L. A., Freyhof, J. & Mehner, T. 2009. Morphological differences between two ecologically similar sympatric fish species. Journal of Fish Biology, 75, 2756-2767. (1.404)
  Kottelat, M. & Freyhof, J. 2009. Notes on the taxonomy and nomenclature of some European freshwater fishes. Ichthyological Exploration of Freshwaters, 20: 75-90. (0.754)
  Schöter, C., Özuluğ, M. & Freyhof, J. 2009. Capoeta caelestis, a new species from River Göksu, Turkey (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 20: 229-236. (0.754)
  Freyhof, J. 2009. Rote Liste der im Süßwasser reproduzierenden Neunaugen und Fische (Cyclostomata & Pisces), fünfte Fassung. Naturschutz und Biologische Vielfalt, 70: 291-316.
  Freyhof; J. & Özuluğ, M. 2009. Pseudophoxinus fahrettini, a new species of spring minnow from Central Anatolia (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 20: 339-346. (0.754)
  Freyhof; J. & Özuluğ, M. 2009. Pseudophoxinus evliyae, a new species of spring minnow from Western Anatolia with remarks on the distribution of P. ninae and the systematic position of P. fahirae (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 20: 347-356. (0.754)
  Lang, N. J., Roe, K. J., Renaud, C. B., Gill, H. S., Potter, I. C., Freyhof, J., Naseka, A., Cochran, P., Pérez, H. E., Habit, E. M., Kuhajda, B. R., Lyons, J., Neely, D. A., Reshtnikov, Y., Salnikov, V., Stoumboudi, M. T. & Mayden, R. L. 2009. Novel relationships among lampreys (Petromyzontiformes) revealed by a taxonomically comprehensive dataset. American Fisheries Society Symposiums, 72: 41-55.
  Borkenhagen, K. & Freyhof, J. 2009. New records of the Levantine endemic cichlid Tristramella simonis (Günther, 1864) from Syria. Cybium, 33: 335-336. (0.541)